EPPO Global Database

Helicoverpa zea(HELIZE)

Hosts

Important note about the classification of host plants in GD:
Categories have been assigned by the EPPO Secretariat on the basis of available data at the time of entry. They correspond to a qualitative evaluation of the importance of the host plant for the pest concerned and remain indicative only.
Further explanation of categories is available in the guide.
Organism Type
Capsicum annuum (CPSAN) Major host
* Hardwick DF (1965) The corn earworm complex. Memoirs of the Entomological Society of Canada, 97, 5–247.
Gossypium hirsutum (GOSHI) Major host
* Johnson MW, Stinner RE, Rabb RL (1975) Ovipositional response of Heliothis zea (Boddie) to its major hosts in North Carolina. Environmental Entomology 4(2):291-297.
Nicotiana tabacum (NIOTA) Major host
* Johnson MW, Stinner RE, Rabb RL (1975) Ovipositional response of Heliothis zea (Boddie) to its major hosts in North Carolina. Environmental Entomology 4(2), 291-297.

* Kolodny-Hirsch DM, Harrison FP (1982) Comparative damage and leaf area consumption by the tobacco budworm and corn earworm on Maryland tobacco. Journal of Economic Entomology, 75(2), 168-172.
Phaseolus vulgaris (PHSVX) Major host
* McLeod P (1988) Seasonal frequency and within-plant distribution of Heliothis zea (Boddie)(Lepidoptera: Noctuidae) eggs on snap bean. Environmental entomology, 17(3), 587-592.
Solanum lycopersicum (LYPES) Major host
* Graham HM, Hernandez Jr NS, Llanes JR (1972) The role of host plants in the dynamics of populations of Heliothis spp. Environmental Entomology 1(4), 424-431.

* Hardwick DF (1965) The corn earworm complex. Memoirs of the Entomological Society of Canada, 97, 5–247.
Solanum melongena (SOLME) Major host
* Hardwick DF (1965) The corn earworm complex. Memoirs of the Entomological Society of Canada, 97, 5–247.
Sorghum bicolor (SORVU) Major host
* Olabimpe O Okosun, Kerry C Allen, James P Glover, Gadi V P Reddy (2021) Biology, Ecology, and Management of Key Sorghum Insect Pests. Journal of Integrated Pest Management 12(1), 4, https://doi.org/10.1093/jipm/pmaa027

* Hardwick DF (1965) The corn earworm complex. Memoirs of the Entomological Society of Canada, 97, 5–247.
Zea mays (ZEAMX) Major host
* Johnson MW, Stinner RE, Rabb RL (1975) Ovipositional response of Heliothis zea (Boddie) to its major hosts in North Carolina. Environmental Entomology 4(2), 291-297.

* Reay-Jones FPF (2019) Pest status and management of corn earworm (Lepidoptera: Noctuidae) in field corn in the United States. Journal of Integrated Pest Management 10(1) 19. https://doi.org/10.1093/jipm/pmz017
Malva parviflora (MALPA) Alternate
* Graham HM, Hernandez Jr NS, Llanes JR (1972) The role of host plants in the dynamics of populations of Heliothis spp. Environmental Entomology 1(4), 424-431.
Abutilon theophrasti (ABUTH) Wild/Weed
* Sudbrink Jr DL, Grant JF (1995) Wild host plants of Helicoverpa zea and Heliothis virescens (Lepidoptera: Noctuidae) in eastern Tennessee. Environmental Entomology 24(5), 1080-1085.
Acalypha persimilis (ACCOS) Wild/Weed
* Allen KC, Elkins BH, Little NS (2024) Acalypha ostryifolia: a natural refuge for Chloridea virescens and Helicoverpa zea (Lepidoptera: Noctuidae) in the Southern United States. Annals of the Entomological Society of America 117(1), 44-48.
------- as Acalypha ostryfolia

* Sudbrink Jr DL, Grant JF (1995) Wild host plants of Helicoverpa zea and Heliothis virescens (Lepidoptera: Noctuidae) in eastern Tennessee. Environmental Entomology 24(5), 1080-1085.
Amaranthus (1AMAG) Wild/Weed
* Kogan M, Helm CG, Kogan J, Brewer E (1989) Distribution and economic importance of Heliothis virescens and Heliothis zea in North, Central, and South America and of their natural enemies and host plants. InProceedings of the Workshop on Biological Control of Heliothis: Increasing the Effectiveness of Natural Enemies, New Delhi, India, 11-15 November 1985 1989. New Delhi, India: Office of International Cooperation & Development, USDA.
Amaranthus palmeri (AMAPA) Wild/Weed
* Kogan M, Helm CG, Kogan J, Brewer E (1989) Distribution and economic importance of Heliothis virescens and Heliothis zea in North, Central, and South America and of their natural enemies and host plants. InProceedings of the Workshop on Biological Control of Heliothis: Increasing the Effectiveness of Natural Enemies, New Delhi, India, 11-15 November 1985 1989. New Delhi, India: Office of International Cooperation & Development, USDA.
Chenopodium (1CHEG) Wild/Weed
* Kogan M, Helm CG, Kogan J, Brewer E (1989) Distribution and economic importance of Heliothis virescens and Heliothis zea in North, Central, and South America and of their natural enemies and host plants. InProceedings of the Workshop on Biological Control of Heliothis: Increasing the Effectiveness of Natural Enemies, New Delhi, India, 11-15 November 1985 1989. New Delhi, India: Office of International Cooperation & Development, USDA.
Cichorium intybus (CICIN) Wild/Weed
* Sudbrink Jr DL, Grant JF (1995) Wild host plants of Helicoverpa zea and Heliothis virescens (Lepidoptera: Noctuidae) in eastern Tennessee. Environmental Entomology 24(5), 1080-1085.
Coronilla varia (CZRVA) Wild/Weed
* Sudbrink Jr DL, Grant JF (1995) Wild host plants of Helicoverpa zea and Heliothis virescens (Lepidoptera: Noctuidae) in eastern Tennessee. Environmental Entomology 24(5), 1080-1085.
Galinsoga quadriradiata (GASCI) Wild/Weed
* Sudbrink Jr DL, Grant JF (1995) Wild host plants of Helicoverpa zea and Heliothis virescens (Lepidoptera: Noctuidae) in eastern Tennessee. Environmental Entomology 24(5), 1080-1085.
------- as G. ciliata.
Geranium dissectum (GERDI) Wild/Weed
* Stadelbacher EA (1981) Role of early-season wild and naturalized host plants in the buildup of the F 1 generation of Heliothis zea and H. virescens in the Delta of Mississippi. Environmental Entomology, 10(5), 766-770.
Glandularia bipinnatifida (VEBBI) Wild/Weed
* Graham HM, Hernandez Jr NS, Llanes JR (1972) The role of host plants in the dynamics of populations of Heliothis spp. Environmental Entomology 1(4), 424-431.
Ipomoea cordatotriloba (IPOTC) Wild/Weed
* Quaintance AL, Brues CT (1905) The cotton bollworm. US Depatment of Agriculture. Bureau of entomology. Bulletin 50. 155 pp.
------- as Ipomoea commutata.
Ipomoea purpurea (PHBPU) Wild/Weed
* Sudbrink Jr DL, Grant JF (1995) Wild host plants of Helicoverpa zea and Heliothis virescens (Lepidoptera: Noctuidae) in eastern Tennessee. Environmental Entomology 24(5), 1080-1085.
Malva neglecta (MALNE) Wild/Weed
* Sudbrink Jr DL, Grant JF (1995) Wild host plants of Helicoverpa zea and Heliothis virescens (Lepidoptera: Noctuidae) in eastern Tennessee. Environmental Entomology 24(5), 1080-1085.
Malva pusilla (MALPU) Wild/Weed
* Quaintance AL, Brues CT (1905) The cotton bollworm. US Depatment of Agriculture. Bureau of entomology. Bulletin 50. 155 pp.
------- as Malva borealis.
Medicago lupulina (MEDLU) Wild/Weed
* Sudbrink Jr DL, Grant JF (1995) Wild host plants of Helicoverpa zea and Heliothis virescens (Lepidoptera: Noctuidae) in eastern Tennessee. Environmental Entomology 24(5), 1080-1085.
Nicotiana repanda (NIORE) Wild/Weed
* Graham HM, Hernandez Jr NS, Llanes JR (1972) The role of host plants in the dynamics of populations of Heliothis spp. Environmental Entomology 1(4), 424-431.
Nuttallanthus canadensis (LINCA) Wild/Weed
* Neunzig HH (1963)  Wild host plants of the corn earworm and the tobacco budworm in eastern North Carolina.  Journal of Economic Entomology, 56(2), 135–139.
Panicum dichotomiflorum (PANDI) Wild/Weed
* Sudbrink Jr DL, Grant JF (1995) Wild host plants of Helicoverpa zea and Heliothis virescens (Lepidoptera: Noctuidae) in eastern Tennessee. Environmental Entomology 24(5), 1080-1085.
Ruellia ciliatiflora (RUELO) Wild/Weed
* Kogan M, Helm CG, Kogan J, Brewer E (1989) Distribution and economic importance of Heliothis virescens and Heliothis zea in North, Central, and South America and of their natural enemies and host plants. InProceedings of the Workshop on Biological Control of Heliothis: Increasing the Effectiveness of Natural Enemies, New Delhi, India, 11-15 November 1985 1989. New Delhi, India: Office of International Cooperation & Development, USDA.
------- as Ruellia lorentziana.
Ruellia runyonii (RUERU) Wild/Weed
* Graham HM, Hernandez Jr NS, Llanes JR (1972) The role of host plants in the dynamics of populations of Heliothis spp. Environmental Entomology 1(4), 424-431.
Sida spinosa (SIDSP) Wild/Weed
* Sudbrink Jr DL, Grant JF (1995) Wild host plants of Helicoverpa zea and Heliothis virescens (Lepidoptera: Noctuidae) in eastern Tennessee. Environmental Entomology 24(5), 1080-1085.
Solanum elaeagnifolium (SOLEL) Wild/Weed
* Graham HM, Hernandez Jr NS, Llanes JR (1972) The role of host plants in the dynamics of populations of Heliothis spp. Environmental Entomology 1(4), 424-431.
Trifolium incarnatum (TRFIN) Wild/Weed
* Stadelbacher EA (1981) Role of early-season wild and naturalized host plants in the buildup of the F 1 generation of Heliothis zea and H. virescens in the Delta of Mississippi. Environmental Entomology, 10(5), 766-770.
Trifolium pratense (TRFPR) Wild/Weed
* Sudbrink Jr DL, Grant JF (1995) Wild host plants of Helicoverpa zea and Heliothis virescens (Lepidoptera: Noctuidae) in eastern Tennessee. Environmental Entomology 24(5), 1080-1085.
Abelmoschus esculentus (ABMES) Host
* Hardwick DF (1965) The corn earworm complex. Memoirs of the Entomological Society of Canada, 97, 5–247.
Apium graveolens (APUGV) Host
* Matthews M (1991) Classification of the Heliothinae. Bulletin of the Natural Resources Institute No. 44. Natural Resources Institute, Chatham, UK.
Arachis hypogaea (ARHHY) Host
* Matthews M (1991) Classification of the Heliothinae. Bulletin of the Natural Resources Institute No. 44. Natural Resources Institute, Chatham, UK.
Avena sativa (AVESA) Host
* Matthews M (1991) Classification of the Heliothinae. Bulletin of the Natural Resources Institute No. 44. Natural Resources Institute, Chatham, UK.
Brassica oleracea (BRSOX) Host
* Quaintance AL, Brues CT (1905) The cotton bollworm. US Depatment of Agriculture. Bureau of entomology. Bulletin 50. 155 pp.
Cajanus cajan (CAJCA) Host
* Lopez Jr JD, Hartstack Jr AW, Beach R (1984) Comparative pattern of emergence of Heliothis zea and H. virescens (Lepidoptera: Noctuidae) from overwintering pupae. Journal of economic entomology 77(6), 1421-1426.
Calendula (1CLDG) Host
* Matthews M (1991) Classification of the Heliothinae. Bulletin of the Natural Resources Institute No. 44. Natural Resources Institute, Chatham, UK.
Cannabis sativa (CNISA) Host
* Britt KE, Kuhar TP, Cranshaw W, McCullough CT, Taylor SV, Arends BR, Burrack H, Pulkoski M, Owens D, Tolosa TA, Zebelo S (2021) Pest management needs and limitations for corn earworm (Lepidoptera: Noctuidae), an emergent key pest of hemp in the United States. Journal of Integrated Pest Management.12(1), 34. https://doi.org/10.1093/jipm/pmab030
Canna indica (CNNIN) Host
* Quaintance AL, Brues CT (1905) The cotton bollworm. US Depatment of Agriculture. Bureau of entomology. Bulletin 50. 155 pp.
Cenchrus americanus (PESGL) Host
* Dourado PM, Pantoja‐Gomez LM, Horikoshi RJ, Carvalho RA, Omoto C, Corrêa AS, Kim JH, Martinelli S, Head GP (2021) Host plant use of Helicoverpa spp. (Lepidoptera: Noctuidae) in the Brazilian agricultural landscape. Pest Management Science 77(2), 780-794.
Chenopodium quinoa (CHEQU) Host
* Chorbadjian RA, Ahumada MI, Urra F, Elgueta M, Gilligan TM (2021) Biogeographical patterns of herbivore arthropods associated with Chenopodium quinoa grown along the Latitudinal Gradient of Chile. Plants 10(12), 2811. doi:10.3390/plants10122811
Cicer arietinum (CIEAR) Host
* Blanco CA, Terán-Vargas AP, López Jr JD, Kauffman JV, Wei X (2007) Densities of Heliothis virescens and Helicoverpa zea (Lepidoptera: Noctuidae) in three plant hosts. Florida Entomologist 90(4), 742-750.
Citrullus lanatus (CITLA) Host
* Quaintance AL, Brues CT (1905) The cotton bollworm. US Depatment of Agriculture. Bureau of entomology. Bulletin 50. 155 pp.
------- as Citrullis vulgaris.
Citrus x aurantium var. sinensis (CIDSI) Host
* Matthews M (1991) Classification of the Heliothinae. Bulletin of the Natural Resources Institute No. 44. Natural Resources Institute, Chatham, UK.
Citrus x limon (CIDLI) Host
* Matthews M (1991) Classification of the Heliothinae. Bulletin of the Natural Resources Institute No. 44. Natural Resources Institute, Chatham, UK.
Cucumis melo (CUMME) Host
* Quaintance AL, Brues CT (1905) The cotton bollworm. US Depatment of Agriculture. Bureau of entomology. Bulletin 50. 155 pp.
Cucumis sativus (CUMSA) Host
* Quaintance AL, Brues CT (1905) The cotton bollworm. US Depatment of Agriculture. Bureau of entomology. Bulletin 50. 155 pp.
Cucurbita pepo (CUUPE) Host
* Quaintance AL, Brues CT (1905) The cotton bollworm. US Depatment of Agriculture. Bureau of entomology. Bulletin 50. 155 pp.
Cynara scolymus (CYUSC) Host
* Kogan M, Helm CG, Kogan J, Brewer E (1989) Distribution and economic importance of Heliothis virescens and Heliothis zea in North, Central, and South America and of their natural enemies and host plants. InProceedings of the Workshop on Biological Control of Heliothis: Increasing the Effectiveness of Natural Enemies, New Delhi, India, 11-15 November 1985 1989. New Delhi, India: Office of International Cooperation & Development, USDA.
Datura stramonium (DATST) Host
* Quaintance AL, Brues CT (1905) The cotton bollworm. US Depatment of Agriculture. Bureau of entomology. Bulletin 50. 155 pp.
Digitaria sanguinalis (DIGSA) Host
* Quaintance AL, Brues CT (1905) The cotton bollworm. US Depatment of Agriculture. Bureau of entomology. Bulletin 50. 155 pp.
------- as Panicum sanguinale.
Erigeron canadensis (ERICA) Host
* Quaintance AL, Brues CT (1905) The cotton bollworm. US Depatment of Agriculture. Bureau of entomology. Bulletin 50. 155 pp.
------ as Erigeron canadense.
Erodium cicutarium (EROCI) Host
* Matthews M (1991) Classification of the Heliothinae. Bulletin of the Natural Resources Institute No. 44. Natural Resources Institute, Chatham, UK.
Fragaria chiloensis (FRACH) Host
* Quaintance AL, Brues CT (1905) The cotton bollworm. US Depatment of Agriculture. Bureau of entomology. Bulletin 50. 155 pp.
Fragaria x ananassa (FRAAN) Host
* Hardwick DF (1965) The corn earworm complex. Memoirs of the Entomological Society of Canada, 97, 5–247.
Gladiolus (1GLAG) Host
* Matthews M (1991) Classification of the Heliothinae. Bulletin of the Natural Resources Institute No. 44. Natural Resources Institute, Chatham, UK.
Glebionis coronaria (CHYCO) Host
* Matthews M (1991) Classification of the Heliothinae. Bulletin of the Natural Resources Institute No. 44. Natural Resources Institute, Chatham, UK.
------- As Chrysanthemum coronarium.
Glycine max (GLXMA) Host
* Hodgson EW, Koch RL, Davis JA, Reisig D, Paula-Moraes SV (2021) Identification and biology of common caterpillars in U.S. soybean. Journal of Integrated Pest Management, 12(1), 13, https://doi.org/10.1093/jipm/pmab006

* Johnson MW, Stinner RE, Rabb RL (1975) Ovipositional response of Heliothis zea (Boddie) to its major hosts in North Carolina. Environmental Entomology 4(2):291-297.

* Swenson SJ, Prishmann-Voldseth DA and Musser FR (2013) Corn earworms (Lepidoptera: Noctuidae) as pests of soybean. Journal of Integrated Pest Management, 4(2), 1-8. https://doi.org/10.1603/IPM13008
Helianthus annuus (HELAN) Host
* Hardwick DF (1965) The corn earworm complex. Memoirs of the Entomological Society of Canada, 97, 5–247.
Hibiscus sabdariffa (HIBSA) Host
* Matthews M (1991) Classification of the Heliothinae. Bulletin of the Natural Resources Institute No. 44. Natural Resources Institute, Chatham, UK.
Lactuca sativa (LACSA) Host
* Hardwick DF (1965) The corn earworm complex. Memoirs of the Entomological Society of Canada, 97, 5–247.
Linum (1LIUG) Host
* Matthews M (1991) Classification of the Heliothinae. Bulletin of the Natural Resources Institute No. 44. Natural Resources Institute, Chatham, UK.
Lonicera japonica (LONJA) Host
* Pair SD (1994) Japanese honeysuckle (Caprifoliaceae): newly discovered host of Heliothis virescens and Helicoverpa zea (Lepidoptera: Noctuidae). Environmental Entomology, 23 (4), 906-911.
Medicago sativa (MEDSA) Host
* Quaintance AL, Brues CT (1905) The cotton bollworm. US Depatment of Agriculture. Bureau of entomology. Bulletin 50. 155 pp.
Oryza sativa (ORYSA) Host
* Matthews M (1991) Classification of the Heliothinae. Bulletin of the Natural Resources Institute No. 44. Natural Resources Institute, Chatham, UK.
Phacelia (1PHCG) Host
* Matthews M (1991) Classification of the Heliothinae. Bulletin of the Natural Resources Institute No. 44. Natural Resources Institute, Chatham, UK.
Phaseolus (1PHSG) Host
* Matthews M (1991) Classification of the Heliothinae. Bulletin of the Natural Resources Institute No. 44. Natural Resources Institute, Chatham, UK.
Phaseolus lunatus (PHSLU) Host
* Matthews M (1991) Classification of the Heliothinae. Bulletin of the Natural Resources Institute No. 44. Natural Resources Institute, Chatham, UK.
Physalis (1PHYG) Host
* Matthews M (1991) Classification of the Heliothinae. Bulletin of the Natural Resources Institute No. 44. Natural Resources Institute, Chatham, UK.
Pisum sativum (PIBSX) Host
* Bimboni H (1977) Ensayos de control químico de la isoca de arveja Helicoverpa zea (Boddie) en cultivo de arveja [Pisum sativum]..[Trials on chemical control of Helicoverpa zea (Boddie) in garden pea crop [Pisum sativum]]. Horticultura. PLAG. IP.-Estación Experimental Agropecuaria San Pedro (Argentina), 15.
Prunus (1PRNG) Host
* Matthews M (1991) Classification of the Heliothinae. Bulletin of the Natural Resources Institute No. 44. Natural Resources Institute, Chatham, UK.
Prunus persica (PRNPS) Host
* Quaintance AL, Brues CT (1905) The cotton bollworm. US Depatment of Agriculture. Bureau of entomology. Bulletin 50. 155 pp.
Prunus sp. (PRNSS) Host
* Quaintance AL, Brues CT (1905) The cotton bollworm. US Depatment of Agriculture. Bureau of entomology. Bulletin 50. 155 pp.
Pyrus communis (PYUCO) Host
* Quaintance AL, Brues CT (1905) The cotton bollworm. US Depatment of Agriculture. Bureau of entomology. Bulletin 50. 155 pp.
Rosa (1ROSG) Host
* Matthews M (1991) Classification of the Heliothinae. Bulletin of the Natural Resources Institute No. 44. Natural Resources Institute, Chatham, UK.
Saccharum officinarum (SACOF) Host
* Matthews M (1991) Classification of the Heliothinae. Bulletin of the Natural Resources Institute No. 44. Natural Resources Institute, Chatham, UK.
Secale cereale (SECCE) Host
* Matthews M (1991) Classification of the Heliothinae. Bulletin of the Natural Resources Institute No. 44. Natural Resources Institute, Chatham, UK.
Setaria italica (SETIT) Host
* Quaintance AL, Brues CT (1905) The cotton bollworm. US Depatment of Agriculture. Bureau of entomology. Bulletin 50. 155 pp.
------- as Chaetochloa italica.
Solanum tuberosum (SOLTU) Host
* Matthews M (1991) Classification of the Heliothinae. Bulletin of the Natural Resources Institute No. 44. Natural Resources Institute, Chatham, UK.
Sphaeralcea sp. (SPHSS) Host
* Matthews M (1991) Classification of the Heliothinae. Bulletin of the Natural Resources Institute No. 44. Natural Resources Institute, Chatham, UK.
-------Sphaeralcea coulteri
Spinacia oleracea (SPQOL) Host
* Matthews M (1991) Classification of the Heliothinae. Bulletin of the Natural Resources Institute No. 44. Natural Resources Institute, Chatham, UK.
Stachys bullata (STABU) Host
* Quaintance AL, Brues CT (1905) The cotton bollworm. US Depatment of Agriculture. Bureau of entomology. Bulletin 50. 155 pp.
------- as Stachys agraria.
Tagetes (1TAGG) Host
* Kogan M, Helm CG, Kogan J, Brewer E (1989) Distribution and economic importance of Heliothis virescens and Heliothis zea in North, Central, and South America and of their natural enemies and host plants. InProceedings of the Workshop on Biological Control of Heliothis: Increasing the Effectiveness of Natural Enemies, New Delhi, India, 11-15 November 1985 1989. New Delhi, India: Office of International Cooperation & Development, USDA.
Trifolium (1TRFG) Host
* Matthews M (1991) Classification of the Heliothinae. Bulletin of the Natural Resources Institute No. 44. Natural Resources Institute, Chatham, UK.
Triticum aestivum (TRZAX) Host
* Matthews M (1991) Classification of the Heliothinae. Bulletin of the Natural Resources Institute No. 44. Natural Resources Institute, Chatham, UK.
Urochloa texana (PANTE) Host
* Quaintance AL, Brues CT (1905) The cotton bollworm. US Depatment of Agriculture. Bureau of entomology. Bulletin 50. 155 pp.
------- as Panicum texanum.
Vicia (1VICG) Host
* Matthews M (1991) Classification of the Heliothinae. Bulletin of the Natural Resources Institute No. 44. Natural Resources Institute, Chatham, UK.
Vicia faba (VICFX) Host
* Hardwick DF (1965) The corn earworm complex. Memoirs of the Entomological Society of Canada, 97, 5–247.
Vigna unguiculata (VIGSI) Host
* Matthews M (1991) Classification of the Heliothinae. Bulletin of the Natural Resources Institute No. 44. Natural Resources Institute, Chatham, UK.
Vitis vinifera (VITVI) Host
* Matthews M (1991) Classification of the Heliothinae. Bulletin of the Natural Resources Institute No. 44. Natural Resources Institute, Chatham, UK.
Xanthium chinense (XANPU) Host
* Kogan M, Helm CG, Kogan J, Brewer E (1989) Distribution and economic importance of Heliothis virescens and Heliothis zea in North, Central, and South America and of their natural enemies and host plants. InProceedings of the Workshop on Biological Control of Heliothis: Increasing the Effectiveness of Natural Enemies, New Delhi, India, 11-15 November 1985 1989. New Delhi, India: Office of International Cooperation & Development, USDA.
------- as Xanthium pennsylvanicum.
Xanthium strumarium (XANST) Host
* Quaintance AL, Brues CT (1905) The cotton bollworm. US Depatment of Agriculture. Bureau of entomology. Bulletin 50. 155 pp.