Important note about the classification of host plants in GD:
Categories have been assigned by the EPPO Secretariat on the basis of available data at the time of entry.
They correspond to a qualitative evaluation of the importance of the host plant for the pest concerned and remain indicative only. Further explanation of categories is available in the guide.
* Athanassiou CG, Phillips TW, Wakil W (2019) Biology and Control of the Khapra Beetle, Trogoderma granarium, a Major Quarantine Threat to Global Food Security. Annual Review of Entomology 64, 131-148. Supplemental Table 1.
------- as peanut
* Lindgren D, Vincent L, Krohne H (1955) The khapra beetle, Trogoderma granarium Everts. Hilgardia 24(1), 1-36.
------- as groundnut
* Global Invasive Species Database (2025) Species profile: Trogoderma granarium. http://www.iucngisd.org/gisd/species.php?sc=142 (27-02-2025).
* Athanassiou CG, Phillips TW, Wakil W (2019) Biology and Control of the Khapra Beetle, Trogoderma granarium, a Major Quarantine Threat to Global Food Security. Annual Review of Entomology 64, 131-148. Supplemental Table 1.
* Global Invasive Species Database (2025) Species profile: Trogoderma granarium. http://www.iucngisd.org/gisd/species.php?sc=142 (27-02-2025).
* Lindgren D, Vincent L, Krohne H (1955) The khapra beetle, Trogoderma granarium Everts. Hilgardia 24(1), 1-36.
* Athanassiou CG, Phillips TW, Wakil W (2019) Biology and Control of the Khapra Beetle, Trogoderma granarium, a Major Quarantine Threat to Global Food Security. Annual Review of Entomology 64, 131-148. Supplemental Table 1.
* Global Invasive Species Database (2025) Species profile: Trogoderma granarium. http://www.iucngisd.org/gisd/species.php?sc=142 (27-02-2025).
* Ciepielewska D, Fornal L, Poplawska M (2000) Infestation of buckwheat grain and its products by pests in storage facilities. Journal of Plant Protection Research 40, 30-34.
* Athanassiou CG, Phillips TW, Wakil W (2019) Biology and Control of the Khapra Beetle, Trogoderma granarium, a Major Quarantine Threat to Global Food Security. Annual Review of Entomology 64, 131-148. Supplemental Table 1.
* Global Invasive Species Database (2025) Species profile: Trogoderma granarium. http://www.iucngisd.org/gisd/species.php?sc=142 (27-02-2025).
* Lindgren D, Vincent L, Krohne H (1955) The khapra beetle, Trogoderma granarium Everts. Hilgardia 24(1), 1-36.
* Global Invasive Species Database (2025) Species profile: Trogoderma granarium. http://www.iucngisd.org/gisd/species.php?sc=142 (27-02-2025).
* Athanassiou CG, Phillips TW, Wakil W (2019) Biology and Control of the Khapra Beetle, Trogoderma granarium, a Major Quarantine Threat to Global Food Security. Annual Review of Entomology 64, 131-148. Supplemental Table 1.
* Lindgren D, Vincent L, Krohne H (1955) The khapra beetle, Trogoderma granarium Everts. Hilgardia 24(1), 1-36.
------- feeding on kernels, citing others
* Global Invasive Species Database (2025) Species profile: Trogoderma granarium. http://www.iucngisd.org/gisd/species.php?sc=142 (27-02-2025).
------- feeding on seed and hay recorded (stored products). Larvae can feed on seeds, but cannot fully develop
* Athanassiou CG, Phillips TW, Wakil W (2019) Biology and Control of the Khapra Beetle, Trogoderma granarium, a Major Quarantine Threat to Global Food Security. Annual Review of Entomology 64, 131-148. Supplemental Table 1.
* Lindgren D, Vincent L, Krohne H (1955) The khapra beetle, Trogoderma granarium Everts. Hilgardia 24(1), 1-36.
* Global Invasive Species Database (2025) Species profile: Trogoderma granarium. http://www.iucngisd.org/gisd/species.php?sc=142 (27-02-2025).
* Athanassiou CG, Phillips TW, Wakil W (2019) Biology and Control of the Khapra Beetle, Trogoderma granarium, a Major Quarantine Threat to Global Food Security. Annual Review of Entomology 64, 131-148. Supplemental Table 1.
* French S, Venette RC (2005) Mini Risk Assessment - Khapra Beetle, Trogoderma granarium (Everts) [Coleoptera: Dermestidae]. CAPS. 22 pp.
------- life cycle not completed on this host (citing others)
* Global Invasive Species Database (2025) Species profile: Trogoderma granarium. http://www.iucngisd.org/gisd/species.php?sc=142 (27-02-2025).
------- feeding host (larvae can feed but cannot fully develop)
* Athanassiou CG, Phillips TW, Wakil W (2019) Biology and Control of the Khapra Beetle, Trogoderma granarium, a Major Quarantine Threat to Global Food Security. Annual Review of Entomology 64, 131-148. Supplemental Table 1.
* Global Invasive Species Database (2025) Species profile: Trogoderma granarium. http://www.iucngisd.org/gisd/species.php?sc=142 (27-02-2025).
* Athanassiou CG, Phillips TW, Wakil W (2019) Biology and Control of the Khapra Beetle, Trogoderma granarium, a Major Quarantine Threat to Global Food Security. Annual Review of Entomology 64, 131-148. Supplemental Table 1.
* Lindgren D, Vincent L, Krohne H (1955) The khapra beetle, Trogoderma granarium Everts. Hilgardia 24(1), 1-36.
* Global Invasive Species Database (2025) Species profile: Trogoderma granarium. http://www.iucngisd.org/gisd/species.php?sc=142 (27-02-2025).
* Okosun OO, Allen KC, Glover JP, Reddy GVP (2021) Biology, Ecology, and Management of Key Sorghum Insect Pests. Journal of Integrated Pest Management 12(1), 1-18.
* Suleiman M, Rugumamu CP (2017) Management of insect pests of stored sorghum using botanicals in Nigerian traditional stores. Journal of Stored Products and Postharvest Research 8(9), 93-102.
* Athanassiou CG, Phillips TW, Wakil W (2019) Biology and Control of the Khapra Beetle, Trogoderma granarium, a Major Quarantine Threat to Global Food Security. Annual Review of Entomology 64, 131-148. Supplemental Table 1.
* Global Invasive Species Database (2025) Species profile: Trogoderma granarium. http://www.iucngisd.org/gisd/species.php?sc=142 (27-02-2025).
* Lindgren D, Vincent L, Krohne H (1955) The khapra beetle, Trogoderma granarium Everts. Hilgardia 24(1), 1-36.
* Athanassiou CG, Phillips TW, Wakil W (2019) Biology and Control of the Khapra Beetle, Trogoderma granarium, a Major Quarantine Threat to Global Food Security. Annual Review of Entomology 64, 131-148. Supplemental Table 1.
* Global Invasive Species Database (2025) Species profile: Trogoderma granarium. http://www.iucngisd.org/gisd/species.php?sc=142 (27-02-2025).
* Athanassiou CG, Phillips TW, Wakil W (2019) Biology and Control of the Khapra Beetle, Trogoderma granarium, a Major Quarantine Threat to Global Food Security. Annual Review of Entomology 64, 131-148. Supplemental Table 1.
* Lindgren D, Vincent L, Krohne H (1955) The khapra beetle, Trogoderma granarium Everts. Hilgardia 24(1), 1-36.
* Global Invasive Species Database (2025) Species profile: Trogoderma granarium. http://www.iucngisd.org/gisd/species.php?sc=142 (27-02-2025).
------- raisins, feeding host (larvae can feed but cannot fully develop)
* Athanassiou CG, Phillips TW, Wakil W (2019) Biology and Control of the Khapra Beetle, Trogoderma granarium, a Major Quarantine Threat to Global Food Security. Annual Review of Entomology 64, 131-148. Supplemental Table 1.
* Lindgren D, Vincent L, Krohne H (1955) The khapra beetle, Trogoderma granarium Everts. Hilgardia 24(1), 1-36.
