EPPO Global Database

Morus alba(MORAL)

Pests

Organism Type
Tetranychus mexicanus (TETRME) Doubtful host
* EPPO (2023) Pest risk analysis for Tetranychus mexicanus. EPPO, Paris
------- uncertain host (see PRA for details)

* Vasconcelos G (2011) Diversidade de ácaros em agroecossistemas e testes para o controle alternativo do ácaro branco, Polyphagotarsonemus latus (Acari, Tarsonemidae) na região de Manaus. Universidade de Sao Paulo.
Aleurocanthus spiniferus (ALECSN) Host
* Cioffi M, Cornara D, Corrado I, Jansen MGM & Porcelli F (2013) The status of Aleurocanthus spiniferus from its unwanted introduction in Italy to date. Bulletin of Insectology 66, 273-281.

* Nugnes F, Laudonia S, Jesu G, Jansen MGM, Bernardo U, Porcelli F (2020) Aleurocanthus spiniferus (Hemiptera: Aleyrodidae) in some European countries: diffusion, hosts, molecular characterization, and natural enemies. Insects 11(1), 42. https://doi.org/10.3390/insects11010042
------- Confirmed host.
Aleuroclava aucubae (TETLAU) Host
* Hári K, Fail J, Streito J.C., Fetykó KG, Szita É, Haltrich A, Vikár D, Radácsi P, Vétek G (2021) First record of Aleuroclava aucubae (Hemiptera Aleyrodidae) in Hungary, with a checklist of whiteflies occurring in the country. Redia 104, 3-7.
Aleurodicus dispersus (ALEDDI) Host
* Boopathi T (2022) New host plants, natural enemy complex and newly distributed potential areas of exotic spiralling whitefly (Hemiptera: Aleyrodidae) in India. Phytoparasitica 50(2), 335-357.
Anoplophora chinensis (ANOLCN) Host
* Lim J, Jung S-Y, Lim J-S, Jang J, Kim K-M, Lee Y-M, Lee B-W (2014) A review of host plants of Cerambycidae (Coleoptera: Chrysomeloidea) with new host records for fourteen Cerambycids, including the Asian longhorn beetle (Anoplophora glabripennis Motschulsky), in Korea. Korean Journal of Applied Entomology 53(2), 111-133.

* Sjöman H, Östberg J & Nilsson J (2014) Review of host trees for the wood-boring pests Anoplophora glabripennis and Anoplophora chinensis: an urban forest perspective. Arboriculture & Urban Forestry 40(3), 143–164.
Anoplophora chinensis (as Morus) (ANOLCN) Host
* Sjöman H, Östberg J & Nilsson J (2014) Review of host trees for the wood-boring pests Anoplophora glabripennis and Anoplophora chinensis: an urban forest perspective. Arboriculture & Urban Forestry 40(3), 143–164.
Anoplophora glabripennis (ANOLGL) Host
* Sjöman H, Östberg J & Nilsson J (2014) Review of host trees for the wood-boring pests Anoplophora glabripennis and Anoplophora chinensis: an urban forest perspective. Arboriculture & Urban Forestry 40(3), 143–164.
Anoplophora glabripennis (as Morus) (ANOLGL) Host
* Lim J, Jung S-Y, Lim J-S, Jang J, Kim K-M, Lee Y-M, Lee B-W (2014) A review of host plants of Cerambycidae (Coleoptera: Chrysomeloidea) with new host records for fourteen Cerambycids, including the Asian longhorn beetle (Anoplophora glabripennis Motschulsky), in Korea. Korean Journal of Applied Entomology 53(2), 111-133.

* Sjöman H, Östberg J & Nilsson J (2014) Review of host trees for the wood-boring pests Anoplophora glabripennis and Anoplophora chinensis: an urban forest perspective. Arboriculture & Urban Forestry 40(3), 143–164.
Apriona cinerea (as Morus) (APRICI) Host
* Singh AP, Bhandari RS, Verma TD (2004) Important insect pests of poplars in agroforestry and strategies for their management in northwestern India. Agroforestry Systems 63, 15–26.
Apriona germari (as Morus) (APRIGE) Host
* Lim J, Jung S-Y, Lim J-S, Jang J, Kim K-M, Lee Y-M, Lee B-W (2014) A review of host plants of Cerambycidae (Coleoptera: Chrysomeloidea) with new host records for fourteen Cerambycids, including the Asian longhorn beetle (Anoplophora glabripennis Motschulsky), in Korea. Korean Journal of Applied Entomology 53(2), 111-133.
Apriona rugicollis (APRIJA) Host
Bactrocera curvipennis (DACUCR) Host
* Leblanc L (2022) The dacine fruit flies (Diptera: Tephritidae: Dacini) of Oceania. Insecta Mundi 0948, 1-167. https://journals.flvc.org/mundi/article/view/131965/135549
Bactrocera dorsalis (DACUDO) Host
* Allwood AJ, Chinajariyawong A, Kritsaneepaiboon S, Drew RAI, Hamacek EL, Hancock DL, Hengsawad C, Jinapin JC, Jirasurat M, Krong CK, Leong CTS, Vijaysegaran S (1999) Host plant records for fruit flies (Diptera: Tephritidae) in Southeast Asia. The Raffles Bulletin of Zoology 47(7), 1-92.
Bactrocera tryoni (DACUTR) Host
* Hancock D, Hamacek EL, Lloyd AC, Elson-Harris MM (2000) The distribution and host plants of fruit flies (Diptera: Tephritidae) in Australia. Queensland Department of Primary Industries, 75 pp.

* Leblanc L, Vueti ET, Drew AI, Allwood AJ (2012) Host plant records for fruit flies (Diptera: Tephritidae: Dacini) in the Pacific islands. Proceedings of the Hawaiian Society 44, 11-53.
'Candidatus Phytoplasma asteris' (PHYPAS) Host
Ceratitis capitata (CERTCA) Host
* Liquido NJ, McQuate GT, Hanlin MA, Suiter KA (2020) Host plants of the Mediterranean fruit fly, Ceratitis capitata (Wiedemann), Version 4.0. Available online at: USDA Compendium of Fruit Fly Host Information (CoFFHI), https://coffhi.cphst.org/
Ceroplastes ceriferus (as Morus) (CERPCE) Host
* Ismailova G (2022) Фитофаги шелковицы в условиях Азербайджана [Mulberry Pests in Azerbaijan Conditions]. Bulletin of Science and Practice 8(10), 54-67..
------- new pest on mulberry. 
Citrus leaf blotch virus (CLBV00) Host
* Xuan Z, Xie J, Yu H, Zhang S, Li R, Cao M (2020) Mulberry (Morus alba) is a new natural host of Citrus leaf blotch virus in China. Plant Disease 105(3), p 716. https://doi.org/10.1094/PDIS-07-20-1580-PDN
------- confirmed host.
Colletotrichum fructicola (COLLFC) Host
* EFSA PLH Panel (EFSA Panel on Plant Health), Bragard C, Dehnen-Schmutz K, Di Serio F, Gonthier P, Jacques M-A, Jaques Miret JA, Justesen AF, MacLeod A, Magnusson CS, Milonas P, Navas-Cortes JA, Parnell S, Potting R, Thulke H-H, Van der Werf W, Civera AV, Yuen J, Zappala L,  Migheli Q, Vloutoglou I, Campese C, Maiorano A, Streissl F, Reignault PL (2021) Scientific Opinion on the pest categorisation of Colletotrichum fructicola. EFSA Journal 19(8), 6803, 41 pp
Diaphania pyloalis (as Morus) (DPHNPY) Host
Dichocrocis punctiferalis (DICHPU) Host
Eutetranychus banksi (EUTEBA) Host
* Migeon A & Dorkeld F (2024) Spider Mites Web: a comprehensive database for the Tetranychidae. Available from https://www1.montpellier.inrae.fr/CBGP/spmweb (Accessed 2024/04/23)
Eutetranychus orientalis (as Morus) (EUTEOR) Host
Euwallacea fornicatus sensu lato (XYLBFO) Host
* Eskalen A, Stouthamer R, Lynch SC, Rugman-Jones PF, Twizeyimana M, Gonzalez A, Thibault T (2013) Host range of Fusarium dieback and its ambrosia beetle (Coleoptera: Scolytinae) vector in southern California. Plant Disease 97(7), 938-951.
Euwallacea fornicatus sensu stricto (EUWAWH) Host
* Mendel Z, Lynch SC, Eskalen A, Protasov A, Maymon M, Freeman S (2021) What determines host range and reproductive performance of an invasive ambrosia beetle Euwallacea fornicatus; lessons from Israel and California. Frontiers in Forests and Global Change 4, 654702.
------- Reproductive host tree in both California (US) and Israel.
Fusarium euwallaceae (FUSAEW) Host
* Eskalen A, Stouthamer R, Lynch SC, Twizeyimana M, Gonzalez A, Thibault T (2013) Host range of Fusarium dieback and its ambrosia beetle (Coleoptera: Scolytinae) vector in southern California. Plant Disease 97(7),  938-951.
Halyomorpha halys (HALYHA) Host
* Mityushev IM (2021) Host plants of Halyomorpha halys in the urban ecosystem on the Azov Sea Coast of Russia. EPPO Bulletin 51(2), 305-310.
------- Abundant on this plant but no significant damage observed in the studied area.
Hishimonus sellatus (HISHSE) Host
* Bella S, Aguin-Pombo D, Aguiar AM, Kunz G, Miralles-Nunez A, Foster S, Sanchez-Garcia I, Wilson MR, D’Urso VE (2022) Discovery of Hishimonus diffractus Dai, Fletcher & Zhang, 2013 (Hemiptera: Auchenorrhyncha: Cicadellidae) in Europe, with remarks on previously recorded species of the genus. Zootaxa 5159(4), 558–570. https://doi.org/10.11646/zootaxa.5159.4.5
Hop stunt viroid (HSVD00) Host
* Lu QY, Ma Y, Han TT, Zhang P, Tang JX, Smith WK, Zhong K, Yu J, Cheng YY, Zhao W (2023) Occurrence and pathogenicity of Hop stunt viroid infecting mulberry (Morus alba) plants in China. Plant Disease 107(early view). https://doi.org/10.1094/PDIS-08-22-1865-RE
------- Natural infection detected in Zhejiang.
Hyphantria cunea (as Morus) (HYPHCU) Host
* Ismailova G (2022) Фитофаги шелковицы в условиях Азербайджана [Mulberry Pests in Azerbaijan Conditions]. Bulletin of Science and Practice 8(10), 54-67.
Icerya aegyptiaca (ICERAE) Host
Icerya seychellarum (ICERSE) Host
* Sánchez-García I, Pérez-Gómez A (2020) First records of Icerya seychellarum [Westwood, 1855] [Hemiptera: Coccomorpha: Monophlebidae] from the iberian peninsula, and new associated species. Revista de la Sociedad Gaditana de Historia Natural 14,  53-55.
Lopholeucaspis japonica (LOPLJA) Host
* Moghaddam M (2013) An annotated checklist of the scale insects of Iran (Hemiptera, Sternorrhyncha, Coccoidea) with new records and distribution data. Zookeys 334, 1-92.
Lycorma delicatula (LYCMDE) Host
* Barringer L, Ciafré CM (2020) Worldwide feeding host plants of spotted lanternfly, with significant additions from North America. Environmental Entomology 49(5), 999–1011.

* Park JD, Kim M-Y, Lee S-G, Shin S-C, Kim J, Park I-K (2009) Biological characteristics of Lycorma delicatula and the control effects of some Insecticides. Korean Journal of Applied Entomology 48(1), 53-57.
------- Nymphs feeding and mild damage were observed in Korea.
Lymantria dispar (as Morus) (LYMADI) Host
* Ismailova G (2022) Фитофаги шелковицы в условиях Азербайджана [Mulberry Pests in Azerbaijan Conditions]. Bulletin of Science and Practice 8(10), 54-67.
Lymantria mathura (LYMAMA) Host
* Roonwal ML (1979) Field-ecological studies on mass eruption, seasonal life-history, nocturnal feeding and activity rhythm, and protective behavior and coloration in the sal defoliator, Lymantria mathura (Lepidoptera: Lymantriidae), in sub-Himalayan forests. Records of the Zoological Survey of India 75, 209-236.
------- A host plant in India.
Maconellicoccus hirsutus (PHENHI) Host
* Martins dos S. D, Fornazier MJ, Peronti ALBG, Culik MP, Souza CAS, Taques RC, Zanuncio Jr JS, Queiroz RB (2019) Maconellicoccus hirsutus (Hemiptera: Pseudococcidae) in Brazil: recent spread, natural enemies, and new hosts. Florida Entomologist 102(2), 438-443.  https://doi.org/10.1653/024.102.0225
------- Confirmed host in Brazil.

* Ülgentürk S, Mohammed EMAM (2016) Scale insects (Hemiptera Coccoidea) on mulberry trees in Turkey. REDIA, XCIX, 2016: 225-228.
------- Confirmed host in Turkey.
Massicus raddei (MALLRA) Host
* Chen SX, Xie YZ, Deng GF (1959) Economic Insect Fauna of China, Fasc. 1. Coleoptera: Cerambycidae. Science Press, Beijing. 43
Meloidogyne enterolobii (MELGMY) Host
* Sun YF, Long HB, Lu FP (2019) First report of the root-knot nematode Meloidogyne enterolobii infecting Mulberry in China. Plant Disease 103, 2481
Meloidogyne enterolobii (as Morus) (MELGMY) Host
* Sun YF, Long HB, Lu FP (2019) First Report of the Root-Knot Nematode Meloidogyne enterolobii Infecting Mulberry in China Plant Disease 103(9), 2481
Mulberry mosaic agent (MUM000) Host
Orgyia leucostigma (as Morus) (HEMELE) Host
* Heppner JB (2003) Lepidoptera of Florida. Part 1. Introduction and catalog. Volume 17 of Arthropods of Florida and neighboring land areas. Division of Plant Industry. Florida Department of Agriculture and Consumer Services. Gainesville, Florida. 670 pp

* Robinson GS, Ackery PR, Kitching IJ, Beccaloni GW & Hernández LM (2010) HOST - A database of the world's Lepidopteran hostplants. Natural History Museum, London. https://www.nhm.ac.uk (Accessed on 7 December 2020 and 21 March 2021)
Orthotospovirus tomatomaculae (TSWV00) Host
* Kil EJ, Chung YJ, Choi HS, Lee S, Kim CS (2020) Life cycle-based host range analysis for Tomato spotted wilt virus in Korea. The Plant Pathology Journal 36(1), 67-75. 
------- Confirmed host.
Parabemisia myricae (PRABMY) Host
Parabemisia myricae (as Morus) (PRABMY) Host
Phryneta leprosa (INESLE) Host
Phymatotrichopsis omnivora (PHMPOM) Host
* Anonymous (1960) Index of Plant Diseases in the United States. Agriculture Handbook no 165, USDA-ARS (US) 531 pp.
Pochazia shantungensis (POCZSH) Host
* Kim DE, Lee H, Kim MJ, Lee DH (2015) Predicting the potential habitat, host plants, and geographical distribution of Pochazia shantungensis (Hemiptera: Ricaniidae) in Korea. Korean Journal of Applied Entomology 54, 179–189.

* Kobayashi S, Suzuki M, Kuwahara R, Park J, Yamada K, Jung S (2024) Reevaluation of taxonomic identity of the recently introduced invasive planthopper, Pochazia shantungensis (Chou & Lu, 1977)(Hemiptera: Fulgoroidea: Ricaniidae) in Japan. Zootaxa 5446(2), 151-178.
Prunus necrotic ringspot virus (PNRSV0) Host
Pseudaulacaspis pentagona (PSEAPE) Host
* Penchevа A, Yovkova M (2016) New data on alien insect pests of ornamental plants in Bulgaria. Forestry ideas, 22, 17-33.
-------- Morus alba f. pendula.

* Stathas GJ, Kartsonas ED, Darras AI, Skouras PJ (2021) Scale insect species (Hemiptera: Coccomorpha) and their natural enemies, recorded on agricultural, ornamental and forest plant species in the wider area of Messenian Province (Peloponnese, Greece), 2000–2020. Hellenic Plant Protection Journal, 14(2), 47-64.
Pseudococcus comstocki (PSECCO) Host
* Pellizzari G, Duso C, Rainato A, Pozzebon A, Zanini G (2012) Phenology, ethology and distribution of Pseudococcus comstocki, an invasive pest in northeastern Italy. Bulletin of Insectology.65(2), 209-215.
Ralstonia pseudosolanacearum (RALSPS) Host
* Pan ZC, Xu J, Prior P, Xu JS, Zhang H, Chen KY, Tian Q, Zhang LQ, Liu L, He LY, Feng J (2013) Development of a specific molecular tool for the detection of epidemiologically active mulberry causing-disease strains of Ralstonia solanacearum phylotype I (historically race 5-biovar 5) in China. European journal of plant pathology 137(2), 377-391.

* She X, He Z, Li H (2018) Genetic structure and phylogenetic relationships of Ralstonia solanacearum strains from diverse origins in Guangdong Province, China. Journal of Phytopathology 166(3):177-186.

* Xu J, Pan ZC, Prior P, Xu JS, Zhang Z, Zhang H, Zhang LQ, He LY, Feng J (2009) Genetic diversity of Ralstonia solanacearum strains from China. European Journal of Pathology 125(4), 641-653.
Ralstonia solanacearum species complex (RALSSO) Host
* Pan ZC, Xu J, Prior P, Xu JS, Zhang H, Chen KY, Tian Q, Zhang LQ, Liu L, He LY, Feng J (2013) Development of a specific molecular tool for the detection of epidemiologically active mulberry causing-disease strains of Ralstonia solanacearum phylotype I (historically race 5-biovar 5) in China. European journal of plant pathology 137(2), 377-391.

* She X, He Z, Li H (2018) Genetic structure and phylogenetic relationships of Ralstonia solanacearum strains from diverse origins in Guangdong Province, China. Journal of Phytopathology 166(3):177-186.

* Xu J, Pan ZC, Prior P, Xu JS, Zhang Z, Zhang H, Zhang LQ, He LY, Feng J (2009) Genetic diversity of Ralstonia solanacearum strains from China. European Journal of Pathology 125(4), 641-653.
Spodoptera eridania (PRODER) Host
* Montezano DG, Specht A, Sosa-Gomez DR, Roque-Specht VF & de Barros NM (2014) Immature stages of Spodoptera eridania (Lepidoptera: Noctuidae): developmental parameters and host plants. Journal of Insect Science 14, 238. https://doi.org/10.1093/jisesa/ieu218
Spodoptera litura (PRODLI) Host
* Ahmad M, Ghaffar A, Rafiq M (2013) Host plants of leaf worm, Spodoptera litura (Fabricius)(Lepidoptera: Noctuidae) in Pakistan. Asian Journal of Agriculture and Biology 1(1), 23-28.
Tetranychus fijiensis (TETRFI) Host
Tobacco streak virus (TSV000) Host
* Rageshwari S, Renukadevi P, Malathi VG, Nakkeeran S (2019) Occurrence of Tobacco streak virus in mulberry. Plant Disease 103(4), p 779.
------- In Tamil Nadu.
Tomato chlorosis virus (TOCV00) Host
* Mamoun Abdel-Salam AM, Rezk AA, Dawoud RA (2019) Biochemical, serological, molecular and natural host studies on Tomato chlorosis virus in Egypt. Pakistan Journal of Biological Sciences 22, 83-94. https://doi.org/10.3923/pjbs.2019.83.126
Xylella fastidiosa (XYLEFA) Host
Xylotrechus namanganensis (XYLONM) Host
* Wang AJ (2001) [Studies on biological characteristics of Xylotrechus namanganensis], Forest Research 14(5), 560–565 (in Chinese).
Zaprionus tuberculatus (ZAPRTU) Host
* Özbek Çatal B, Çalişkan Keçe AF, Ulusoy MR (2021) Distribution and host plants of Drosophilidae (Diptera) species detected in fruit orchards of the Eastern Mediterranean Region of Turkey. Mustafa Kemal University Journal of Agricultural Sciences 26(2), 431-442. https://doi.org/10.37908/mkutbd.873838
Apriona germari (APRIGE) Major host
* Lim J, Jung S-Y, Lim J-S, Jang J, Kim K-M, Lee Y-M, Lee B-W (2014) A review of host plants of Cerambycidae (Coleoptera: Chrysomeloidea) with new host records for fourteen Cerambycids, including the Asian longhorn beetle (Anoplophora glabripennis Motschulsky), in Korea. Korean Journal of Applied Entomology 53(2), 111-133.
Diaphania pyloalis (DPHNPY) Major host
Hyphantria cunea (HYPHCU) Major host
Meloidogyne mali (MELGMA) Major host
* Toida Y (1986) On the cultural control of the mulberry attacking root-knot nematode, Meloidogyne mali Itoh, Ohshima et Ichinohe. Sansi-kenkyu (Acta Sericologica), No. 136,Jan. 1986 [text, not figures – 2058 words]
------- confirmed host
Peridiopsora mori (AECIMO) Major host
Psacothea hilaris (PSACHI) Major host
* Lim J, Jung S-Y, Lim J-S, Jang J, Kim K-M, Lee Y-M, Lee B-W (2014) A review of host plants of Cerambycidae (Coleoptera: Chrysomeloidea) with new host records for fourteen Cerambycids, including the Asian longhorn beetle (Anoplophora glabripennis Motschulsky), in Korea. Korean Journal of Applied Entomology 53(2), 111-133.

* Lupi D, Jucker C, Colombo M (2013) Distribution and biology of the yellow-spotted longicorn beetle Psacothea hilaris hilaris (Pascoe) in Italy. EPPO Bulletin, 43, 316–322.
------- Rare host in Italy (compared to Ficus carica).
Psacothea hilaris (as Morus) (PSACHI) Major host
* Lim J, Jung S-Y, Lim J-S, Jang J, Kim K-M, Lee Y-M, Lee B-W (2014) A review of host plants of Cerambycidae (Coleoptera: Chrysomeloidea) with new host records for fourteen Cerambycids, including the Asian longhorn beetle (Anoplophora glabripennis Motschulsky), in Korea. Korean Journal of Applied Entomology 53(2), 111-133.
Trichoferus campestris (as Morus) (HESOCA) Major host
* CAPS (2019) Trichoferus campestris. Cooperative Agricultural Pest Survey pest datasheets. http://download.ceris.purdue.edu/file/3869
------- Preferred living host.
Xylotrechus chinensis (XYLOCH) Major host
* Han Y, Lee D (2010) Taxonomic review of the genus Xylotrechus (Coleoptera: Cerambycidae: Cerambycinae) in Korea with a newly recorded species. Korean Journal of Applied Entomology 49(2), 69–82. https://doi.org/10.5656/KSAE.2010.49.2.069

* Lin M, Ge S, Xiao N (2021) A study of the genus Xylotrechus Chevrolat (Coleoptera: Cerambycidae) from Beijing, China. Entomotaxonomia 43(3), 20 pp. https://doi.org/10.11680/entomotax.2021021

* Sarto i Monteys V, Torras i Tutusaus G (2018) A new alien invasive longhorn beetle, Xylotrechus chinensis (Cerambycidae), is infesting mulberries in Catalonia (Spain). Insects 9(2), 52. https://doi.org/10.3390/insects9020052
Xylotrechus chinensis (as Morus) (XYLOCH) Major host
* Han Y, Lee D (2010) Taxonomic review of the genus Xylotrechus (Coleoptera: Cerambycidae: Cerambycinae) in Korea with a newly recorded species. Korean Journal of Applied Entomology 49(2), 69–82. https://doi.org/10.5656/KSAE.2010.49.2.069

* Sarto i Monteys V, Torras i Tutusaus G (2018) A new alien invasive longhorn beetle, Xylotrechus chinensis (Cerambycidae), is infesting mulberries in Catalonia (Spain). Insects 9(2), 52. https://doi.org/10.3390/insects9020052
Icerya purchasi (as Morus) (ICERPU) Wild/Weed
* Ismailova G (2022) Фитофаги шелковицы в условиях Азербайджана [Mulberry Pests in Azerbaijan Conditions]. Bulletin of Science and Practice 8(10), 54-67.