* Matousek J, Siglova K, Jakse J, Radisek S, Brass JRJ, Tsushima T, Gucek T, Duraisamy GS, Sano T, Steger G (2017)
Propagation and some physiological effects of Citrus bark cracking viroid and Apple fruit crinkle viroid in multiple infected hop (Humulus lupulus L.). Journal of Plant Physiology 213, 166-177.
* Suzuki T, Fujibayashi M, Hataya T, Taneda A, He YH, Tsushima T, Duraisamy GS, Siglova K, Matousek J, Sano T (2017) Characterization of host-dependent mutations of apple fruit crinkle viroid replicating in newly identified experimental hosts suggests maintenance of stem-loop structures in the left-hand half of the molecule is important for replication. Journal of General Virology 98(3), 506-516.
------- Cultivated hop (natural host) and wild hop (experimental host).
* Jakse J, Radisek S, Pokorn T, Matousek J, Javornik B (2015) Deep-sequencing revealed Citrus bark cracking viroid (CBCVd) as a highly aggressive pathogen on hop. Plant Patholology, 64: 831–842. doi:10.1111/ppa.1232
------- confirmed host
*Marquez‐Molins J, Gomez G, Pallas V (2020) Hop stunt viroid: A polyphagous pathogenic RNA that has shed light on viroid–host interactions. Molecular Plant Pathology. https://doi.org/10.1111/mpp.13022
* Sasaki M, Shikata E (1977) On some properties of hop stunt disease agent, a viroid. Proceedings of the Japan Academy. Series B, Physical and Biological Sciences, 53, 109–112.